Балегари

Балегари
	Средњоевропски балегари
Научна класификација
Домен:	Eukaryota
Царство:	Animalia
Тип:	Arthropoda
Класа:	Insecta
Ред:	Coleoptera
Натпородица:	Scarabaeoidea
Породица:	Scarabaeidae; Latreille, 1802
Потпородице
	Aclopinae; Aegialiinae; Allidiostomatinae; Aphodiinae; Cetoniinae; Chironinae; Dynastinae; Dynamopodinae; Eremazinae; Euchirinae; Melolonthinae; Orphninae; Pachypodinae; Phaenomeridinae; Phileurinae; Rutelinae; Scarabaeinae; Termitotroginae; Cretoscarabaeinae †; Lithoscarabaeinae †; Prototroginae †;

Балегари^[1] или котрљани^[1] (лат. Scarabaeidae) су породица инсеката из реда тврдокрилаца (Coleoptera). Познато је преко 28.000 врста, од којих се већина храни биљкама.^[1] Док се већина врста из потпородице правих балегара Scarabaeinae, која укључује више од 5.000 врста,^[2] храни балегом. Највећи број врста из ове породице је присутан у тропима.^[1]

Опис

Балегари су здепасти инсекти, који достижу од 1,5 до 160 mm, зависно од врсте. Јављају се у различитим бојама, од црне до металик зелено-плаве или бакарне. Имају карактеристичне антене у облику палице, које могу склупчати или испружити да би осетили мирисе. Предње ноге многих врста су широке и прилагођене копању. Мужјаци неких врста (код неких врста и женке) имају рогове на глави или на вратном штиту (пронотуму), које користе у борби око женки или хране.^[3]

Митологија

Стари Египћани су за врсту балегара скарабеј (свети скарабеј, котрљан или балегар) (лат. Scarabaeus sacer) веровали да је света. У староегипатској религији балегарово котрљање куглице балеге по песку била је метафора за пут сунца по небу. Египатским амајлијама које су представљале светог скарабеја се трговало по целом Средоземљу.^[3]

Систематика

Родови

Acoma Casey, 1889
Acrossus Mulsant, 1842
Adoretus Dejean, 1833
Aegialia Latreille, 1807
Agoliinus Schmidt, 1913
Agrilinellus Dellacasa, Dellacasa and Gordon, 2008
Aidophus Balthasar, 1963
Alloblackburneus Bordat, 2009
Amblonoxia Reitter, 1902
Amphimallon Latreille, 1825
Ancognatha Erichson, 1847
Annegialia Howden, 1971
Anomala Samouelle, 1819
Anomalacra Casey, 1915
Anoplognatho Rivers, 1888
Apeltastes Howden, 1968
Aphodius Illiger, 1798
Aphonus LeConte, 1856
Aphotaenius Cartwright, 1952
Archophileurus Kolbe, 1910
Arctodium Burmeister, 1844
Ataeniopsis Petrovitz, 1973
Ataenius Harold, 1867
Ateuchus Weber, 1801
Australaphodius Balthasar, 1942
Ballucus Gordon and Skelley, 2007
Blackburneus Schmidt, 1913
Caelius Lewis, 1895
Calamosternus Motschulsky, 1859
Caligodorus Gordon and Skelley, 2007
Calomacraspis Bates, 1888
Canthidium Erichson, 1847
Canthon Hoffmannsegg, 1817
Centrochilus Krikken, 1976
Cephalocyclus Dellacasa, Gordon and Dellacasa, 1998
Chaunocolus Saylor, 1937
Chilothorax Motschulsky, 1859
Chlorixanthe Bates, 1889
Chnaunanthus Burmeister, 1844
Chrysina Kirby, 1828
Cinacanthus Schmidt, 1913
Coelotrachelus Schmidt, 1913
Coenonycha Horn, 1876
Colobopterus Mulsant, 1842
Copris Geoffroy, 1762
Coprophanaeus D'Olsoufieff, 1924
Coscinocephalus Prell, 1936
Cotalpa Burmeister, 1844
Cotinis Burmeister, 1842
Cremastocheilus Knoch, 1801
Cryptoscatomaseter Gordon and Skelley, 2007
Cyclocephala Dejean, 1821
Dellacasiellus Gordon and Skelley, 2007
Deltochilum Eschscholtz, 1822
Dialytellus Brown, 1929
Dialytes Harold, 1869
Dialytodius Gordon and Skelley, 2007
Diapterna Horn, 1887
Diastictus Mulsant, 1842
Dichelonyx Harris, 1827
Dichotomius Hope, 1838
Dinacoma Casey, 1889
Diplotaxis Kirby, 1837
Drepanacanthoides Schmidt, 1913
Dynastes MacLeay, 1819
Dyscinetus Harold, 1869
Ectinoplectron Ohaus, 1915
Enema Hope, 1837
Euetheola Bates, 1888
Euoniticellus Janssens, 1953
Euparia LePeletier and Serville, 1828
Euparixia Brown, 1927
Euphoria Burmeister, 1842
Eupleurus Mulsant, 1842
Flaviellus Gordon and Skelley, 2007
Fossocarus Howden, 1961
Genuchinus Westwood, 1874
Geomyphilus Gordon and Skelley, 2007
Geopsammodius Gordon and Pittino, 1992
Gillaspytes Howden, 1980
Gnorimella Casey, 1915
Golofa Hope, 1837
Gronocarus Schaeffer, 1927
Gymnetina Casey, 1915
Gymnetis MacLeay, 1819
Gymnopyge Linell, 1896
Halffterinetis Morón and Nogueira, 2007
Haroldiataenius Chalumeau, 1981
Haroldiellus Gordon and Skelley, 2007
Hemiphileurus Kolbe, 1910
Hologymnetis Martínez, 1949
Homoiosternus Ohaus, 1901
Hoplia Illiger, 1803
Hornietus Stebnicka, 2000
Hypothyce Howden, 1968
Hypotrichia LeConte, 1861
Inca LePeletier and Serville, 1828
Irrasinus Gordon and Skelley, 2007
Isonychus Mannerheim, 1829
Labarrus Mulsant and Rey, 1870
Lechorodius Gordon and Skelley, 2007
Leiopsammodius Rakovic, 1981
Leptohoplia Saylor, 1935
Liatongus Reitter, 1893
Lichnia Erichson, 1835
Liothorax Motschulsky, 1859
Lissomelas Bates, 1889
Luxolinus Gordon and Skelley, 2007
Macrodactylus Dejean, 1821
Maculaphodius Gordon and Skelley, 2007
Maladera Mulsant and Rey, 1871
Malagoniella Martínez, 1961
Martineziana Chalumeau and Ozdikmen, 2006
Megasoma Kirby, 1825
Melanocanthon Halffter, 1958
Melinopterus Mulsant, 1842
Mendidius Harold, 1868
Merogyrus Gordon and Skelley, 2007
Micraegialia Brown, 1931
Nefoncerus Saylor, 1938
Neopsammodius Rakovic, 1986
Neotrichonotulus Dellacasa, Gordon and Dellacasa, 2004
Nialaphodius Kolbe, 1908
Nipponoserica Nomura, 1973
Odontolytes Koshantschikov, 1916
Odontopsammodius Gordon and Pittino, 1992
Oncerus LeConte, 1856
Oniticellus LePeletier and Serville, 1828
Onitis Fabricius, 1798
Ontherus Erichson, 1847
Onthophagus Latreille, 1802
Orizabus Fairmaire, 1878
Oscarinus Gordon and Skelley, 2007
Osmoderma LePeletier and Serville, 1828
Otophorus Mulsant, 1842
Oxygrylius Casey, 1915
Oxyomus Dejean, 1833
Parabyrsopolis Ohaus, 1915
Parachrysina Bates, 1888
Paracotalpa Ohaus, 1915
Parapsammodius Gordon and Pittino, 1992
Parastasia Westwood, 1842
Parataenius Balthasar, 1961
Pardalosus Gordon and Skelley, 2007
Pelidnota MacLeay, 1819
Phaeaphodius Reitter, 1892
Phanaeus MacLeay, 1819
Phileurus Latreille, 1807
Phobetus LeConte, 1856
Phyllophaga Harris, 1827
Planolinellus Dellacasa and Dellacasa, 2005
Planolinoides Dellacasa and Dellacasa, 2005
Planolinus Mulsant and Rey, 1870
Platytomus Mulsant, 1842
Plectris LePeletier and Serville, 1828
Plectrodes Horn, 1867
Plesiosternus Morón, 1983
Pleurophorus Mulsant, 1842
Podolasia Harold, 1869
Podostena Howden, 1997
Polyphylla Harris, 1841
Popillia Dejean, 1821
Protaetia Burmeister, 1842
Psammodius Fallén, 1807
Pseudagolius Schmidt, 1913
Pseudataenius Brown, 1927
Pseudocanthon Bates, 1887
Pseudocotalpa Hardy, 1971
Psilocnemis Burmeister, 1842
Rhysothorax Bedel, 1911
Rhyssemus Mulsant, 1842
Rugaphodius Gordon and Skelley, 2007
Rutela Latreille, 1802
Rutelisca Bates, 1888
Saprosites Redtenbacher, 1858
Scabrostomus Gordon and Skelley, 2007
Schaefferellus Gordon and Skelley, 2007
Serica MacLeay, 1819
Setodius Gordon and Skelley, 2007
Sisyphus Latreille, 1807
Stenotothorax Schmidt, 1913
Strategus Kirby, 1828
Strigoderma Burmeister, 1844
Strigodius Gordon and Skelley, 2007
Tesarius Rakovic, 1981
Tetraclipeoides Schmidt, 1913
Teuchestes Mulsant, 1842
Thyce LeConte, 1856
Tomarus Erichson, 1847
Trichiorhyssemus Clouët, 1901
Trichiotinus Casey, 1915
Trichonotulus Bedel, 1911
Trigonopeltastes Burmeister, 1840
Uroxys Westwood, 1842
Valgus Scriba, 1790
Warwickia Smith and Evans, 2005
Xeropsamobeus Saylor, 1937
Xyloryctes Hope, 1837

Врсте

Acoma arizonica Brown, 1929
Acoma brunnea Casey, 1889
Acoma cazieri Saylor, 1948
Acoma confusa Van Dyke, 1928
Acoma conjuncta Howden, 1962
Acoma dilemma Saylor, 1948
Acoma diminiata Howden, 1958
Acoma evansi Howden, 1962
Acoma gibsoni Howden, 1962
Acoma glabrata Cazier, 1953
Acoma granulifrons Howden, 1958
Acoma incognita Howden, 1958
Acoma knulli Howden, 1958
Acoma leechi Cazier, 1953
Acoma martini Howden, 1962
Acoma mimica Howden, 1962
Acoma minuta Cazier, 1953
Acoma mixta Howden, 1958
Acoma ochlera Howden, 1958
Acoma parva Howden, 1958
Acoma robusta Van Dyke, 1928
Acoma rossi Saylor, 1948
Acoma rufula Howden, 1958
Acoma seticollis Howden, 1958
Acoma sexfoliata Saylor, 1948
Acoma stathamae Cazier, 1953
Acrossus rubripennis (Horn, 1870)
Acrossus rufipes (Linnaeus, 1758)
Adoretus compressus (Weber, 1801)
Adoretus sinicus Burmeister, 1855
Aegialia amplipunctata Gordon and Cartwright, 1988
Aegialia arenaria (Fabricius, 1787)
Aegialia blanchardi Horn, 1887
Aegialia carri Gordon and Cartwright, 1988
Aegialia cartwrighti Stebnicka, 1977
Aegialia concinna Gordon and Cartwright, 1977
Aegialia conferta Horn, 1871
Aegialia convexa Fall, 1932
Aegialia crassa LeConte, 1857
Aegialia crescenta Gordon and Cartwright, 1977
Aegialia criddlei Brown, 1931
Aegialia cylindrica (Eschscholtz, 1822)
Aegialia exarata Mannerheim, 1853
Aegialia hardyi Gordon and Cartwright, 1977
Aegialia kelsoi Gordon and Cartwright, 1988
Aegialia knighti Gordon and Rust, 1997
Aegialia lacustris LeConte, 1850
Aegialia latispina LeConte, 1878
Aegialia magnifica Gordon and Cartwright, 1977
Aegialia mcclevei Gordon, 1990
Aegialia nana Brown, 1931
Aegialia nigrella Brown, 1931
Aegialia opaca Brown, 1931
Aegialia opifex Horn, 1887
Aegialia punctata Brown, 1931
Aegialia spinosa Gordon and Cartwright, 1988
Aegialia terminalis Brown, 1931
Agoliinus albertanus (Brown, 1928)
Agoliinus aleutus (Eschscholtz, 1822)
Agoliinus anthracus Gordon and Skelley, 2007
Agoliinus aquilonarius (Brown, 1928)
Agoliinus ashworthi (Gordon, 2006)
Agoliinus bidentatus (Schmidt, 1907)
Agoliinus canadensis (Garnett, 1920)
Agoliinus congregatus (Mannerheim, 1853)
Agoliinus corruptor (Brown, 1929)
Agoliinus cruentatus (LeConte, 1878)
Agoliinus explanatus (LeConte, 1878)
Agoliinus guttatus (Eschscholtz, 1823)
Agoliinus hatchi Gordon and Skelley, 2007
Agoliinus idahoensis Gordon and Skelley, 2007
Agoliinus incommunis (Fall, 1932)
Agoliinus ingenursus Gordon and Skelley, 2007
Agoliinus leopardus (Horn, 1870)
Agoliinus malkini (Hatch, 1971)
Agoliinus manitobensis (Brown, 1928)
Agoliinus piceatus (Robinson, 1946)
Agoliinus plutonicus (Fall, 1907)
Agoliinus poudreus Gordon and Skelley, 2007
Agoliinus praealtus Gordon and Skelley, 2007
Agoliinus sigmoideus (Van Dyke, 1918)
Agoliinus wickhami (Brown, 1928)
Agrilinellus nuriae Dellacasa, Dellacasa and Gordon, 2008
Agrilinellus ornatus (Schmidt, 1911)
Aidophus parcus (Horn, 1887)
Aidophus skelleyi Harpootlian and Gordon, 2002
Alloblackburneus aegrotus (Horn, 1870)
Alloblackburneus cavidomus (Brown, 1927)
Alloblackburneus cynomysi (Brown, 1927)
Alloblackburneus fordi (Gordon, 1974)
Alloblackburneus geomysi (Cartwright, 1939)
Alloblackburneus guadalajarae M. Dellacasa, G. Dellacasa and Gordon, 2011
Alloblackburneus ibanezbernali M. Dellacasa, G. Dellacasa and Gordon, 2011
Alloblackburneus lentus (Horn, 1870)
Alloblackburneus rubeolus (Palisot de Beauvois, 1809)
Alloblackburneus saylori (Hinton, 1934)
Alloblackburneus tenuistriatus (Horn, 1887)
Alloblackburneus troglodytes (Hubbard, 1894)
Amblonoxia bidentata (Fall, 1932)
Amblonoxia carpenteri (LeConte, 1876)
Amblonoxia fieldi (Fall, 1908)
Amblonoxia harfordi (Casey, 1889)
Amblonoxia palpalis (Horn, 1880)
Amblonoxia riversi (Casey, 1895)
Amphimallon majale (Razumowski, 1789)
Ancognatha manca (LeConte, 1866)
Annegialia ataeniformis Howden, 1971
Anomala adscita (Robinson, 1941)
Anomala antennata Schaeffer, 1906
Anomala arida Casey, 1915
Anomala beckeri Ohaus, 1897
Anomala binotata (Gyllenhal, 1817)
Anomala butleri (Howden, 1955)
Anomala carinifrons Bates, 1888
Anomala carlsoni Hardy, 1976
Anomala castaniceps Bates, 1888
Anomala cavifrons LeConte, 1867
Anomala centralis LeConte, 1863
Anomala chevrolati Bates, 1888
Anomala cincta Say, 1835
Anomala crinicollis Ohaus, 1902
Anomala delicata Casey, 1915
Anomala diabla Potts, 1976
Anomala digressa Casey, 1915
Anomala discoidalis Bates, 1888
Anomala dubia (Scopoli, 1763)
Anomala ellipsis Casey, 1915
Anomala exigua (Schwarz, 1878)
Anomala eximia Potts, 1976
Anomala flavilla Bates, 1888
Anomala flavipennis Burmeister, 1844
Anomala flohri Ohaus, 1897
Anomala foraminosa Bates, 1888
Anomala hardyorum Potts, 1976
Anomala histrionella Bates, 1888
Anomala imperialae Potts, 1976
Anomala innuba (Fabricius, 1787)
Anomala insitiva Robinson, 1938
Anomala kanei Potts, 1976
Anomala lucicola (Fabricius, 1798)
Anomala ludoviciana Schaeffer, 1906
Anomala marginata (Fabricius, 1793)
Anomala mendica Casey, 1915
Anomala nimbosa Casey, 1915
Anomala oblivia Horn, 1884
Anomala orientalis (Waterhouse, 1875)
Anomala parvula Burmeister, 1844
Anomala peninsularis Schaeffer, 1906
Anomala robinsoni Potts, 1974
Anomala sabinae Potts, 1976
Anomala semilivida LeConte, 1878
Anomala suavis Potts, 1976
Anomala sulcatula Burmeister, 1844
Anomala tibialis Schaeffer, 1906
Anomala umbra Casey, 1915
Anomala undulata Melsheimer, 1845
Anomalacra clypealis (Schaeffer, 1907)
Anoplognatho dunnianus Rivers, 1888
Apeltastes elongatus Howden, 1968
Aphodius fimetarius (Linnaeus, 1758)
Aphonus brevicruris Cartwright, 1944
Aphonus castaneus (Melsheimer, 1845)
Aphonus densicauda Casey, 1915
Aphonus texanus Gill and Howden, 1985
Aphonus tridentatus (Say, 1823)
Aphonus variolosus (LeConte, 1847)
Aphotaenius carolinus (Van Dyke, 1928)
Aphotaenius howdeni Cartwright, 1963
Archophileurus cribrosus (LeConte, 1854)
Arctodium discolor (Erichson, 1835)
Arctodium mahdii Hawkins, 2006
Arctodium planum (Blanchard, 1850)
Arctodium vulpinum (Erichson, 1835)
Ataeniopsis duncani (Cartwright, 1974)
Ataeniopsis edistoi (Cartwright, 1974)
Ataeniopsis figurator (Harold, 1874)
Ataeniopsis parkeri (Cartwright, 1974)
Ataeniopsis rugopygus (Cartwright, 1974)
Ataeniopsis saxatilis (Cartwright, 1944)
Ataenius abditus (Haldeman, 1848)
Ataenius aequalis Harold, 1880
Ataenius alternatus (Melsheimer, 1845)
Ataenius apicalis Hinton, 1937
Ataenius barberi Cartwright, 1974
Ataenius brevicollis (Wollaston, 1854)
Ataenius brevis Fall, 1930
Ataenius californicus Horn, 1887
Ataenius carinator Harold, 1874
Ataenius castaniellus Bates, 1887
Ataenius cognatus (LeConte, 1858)
Ataenius confertus Fall, 1909
Ataenius crenulatus Schmidt, 1910
Ataenius cribrithorax Bates, 1887
Ataenius cylindrus Horn, 1871
Ataenius desertus Horn, 1871
Ataenius ecruensis Stebnicka and Lago, 2005
Ataenius erratus Fall, 1930
Ataenius exiguus Brown, 1932
Ataenius fattigi Cartwright, 1948
Ataenius glaseri Cartwright, 1974
Ataenius gracilis (Melsheimer, 1845)
Ataenius heinekeni (Wollaston, 1854)
Ataenius hesperius Cartwright, 1974
Ataenius hirsutus Horn, 1871
Ataenius imbricatus (Melsheimer, 1845)
Ataenius inquisitus Horn, 1887
Ataenius insculptus Horn, 1887
Ataenius languidus Schmidt, 1911
Ataenius liogaster Bates, 1887
Ataenius lobatus Horn, 1871
Ataenius miamii Cartwright, 1934
Ataenius oklahomensis Brown, 1930
Ataenius opatrinus Harold, 1867
Ataenius ovatulus Horn, 1871
Ataenius pacificus (Sharp, 1879)
Ataenius peregrinator Harold, 1877
Ataenius picinus Harold, 1867
Ataenius platensis (Blanchard, 1847)
Ataenius puncticollis LeConte, 1868
Ataenius robustus Horn, 1871
Ataenius scabrelloides Petrovitz, 1962
Ataenius scabrellus Schmidt, 1909
Ataenius sculptor Harold, 1868
Ataenius setiger Bates, 1887
Ataenius spretulus (Haldeman, 1848)
Ataenius stephani Cartwright, 1974
Ataenius stercorator (Fabricius, 1775)
Ataenius strigatus (Say, 1823)
Ataenius texanus Harold, 1874
Ataenius utahensis Cartwright, 1974
Ataenius vandykei Cartwright, 1974
Ataenius waltherhorni Balthasar, 1938
Ataenius wenzelii Horn, 1887
Ateuchus floridensis Génier, 2000
Ateuchus histeroides Weber, 1801
Ateuchus illaesum (Harold, 1868)
Ateuchus lecontei (Harold, 1868)
Ateuchus rodriguezi (Preudhomme de Borre, 1886)
Ateuchus texanus (Robinson, 1948)
Australaphodius frenchi (Blackburn, 1892)
Ballucus barri (Gordon, 2006)
Ballucus bruneaui Gordon and Skelley, 2007
Ballucus obtusus (LeConte, 1878)
Ballucus oregonus Gordon and Skelley, 2007
Blackburneus amazonicus M. Dellacasa, G. Dellacasa and Gordon, 2011
Blackburneus argentinensis (Schmidt, 1909)
Blackburneus caracaensis (Petrovitz, 1970)
Blackburneus charmionus (Bates, 1887)
Blackburneus diminutus (Bates, 1887)
Blackburneus erythrinus (Bates, 1887)
Blackburneus furcatus (Schmidt, 1909)
Blackburneus guatemalensis (Bates, 1887)
Blackburneus indio (Petrovitz, 1973)
Blackburneus laxepunctatus (Schmidt, 1910)
Blackburneus richteri (Schmidt, 1911)
Blackburneus sanfilippoi M. Dellacasa, G. Dellacasa and Gordon, 2011
Blackburneus saylorea (Robinson, 1940)
Blackburneus stercorosus (Melsheimer, 1844)
Blackburneus surinamensis M. Dellacasa, G. Dellacasa and Gordon, 2011
Blackburneus teposcolulaensis M. Dellacasa, G. Dellacasa and Gordon, 2011
Blackburneus thomasi M. Dellacasa, G. Dellacasa and Gordon, 2011
Blackburneus xanthus (Bates, 1887)
Caelius browni (Saylor, 1934)
Caelius humeralis (Brown, 1931)
Caelius montana (Brown, 1931)
Caelius rufescens (Horn, 1887)
Calamosternus granarius (Linnaeus, 1767)
Caligodorus opacus (LeConte, 1872)
Caligodorus tuberosus (Barrett, 1932)
Caligodorus vandykei (Barrett, 1931)
Calomacraspis splendens (Burmeister, 1844)
Canthidium laetum Harold, 1867
Canthidium macclevei Kohlmann and Solís, 2006
Canthidium pseudopuncticolle Solís and Kohlmann, 2004
Canthon bisignatus Balthasar, 1939
Canthon chalcites (Haldeman, 1843)
Canthon coahuilensis Howden, 1966
Canthon cyanellus LeConte, 1859
Canthon depressipennis LeConte, 1859
Canthon ebenus (Say, 1823)
Canthon femoralis (Chevrolat, 1834)
Canthon forreri Bates, 1887
Canthon humectus (Say, 1832)
Canthon imitator Brown, 1946
Canthon indigaceus LeConte, 1866
Canthon lecontei Harold, 1868
Canthon melanus Robinson, 1948
Canthon mixtus Robinson, 1948
Canthon obliquus Horn, 1894
Canthon pilularius (Linnaeus, 1758)
Canthon praticola LeConte, 1859
Canthon probus (Germar, 1824)
Canthon puncticollis LeConte, 1866
Canthon simplex LeConte, 1857
Canthon vigilans LeConte, 1858
Canthon viridis (Palisot de Beauvois, 1805)
Centrochilus howdeni Krikken, 1976
Cephalocyclus bordati Dellacasa, Dellacasa and Gordon, 2007
Cephalocyclus carmenae Dellacasa, Dellacasa and Gordon, 2007
Cephalocyclus durangoensis (Bates, 1887)
Cephalocyclus fuliginosus (Harold, 1863)
Cephalocyclus gravidus (Harold, 1863)
Cephalocyclus hogei (Bates, 1887)
Cephalocyclus howdenorum Dellacasa, Dellacasa and Gordon, 2007
Cephalocyclus lagoi Dellacasa, Dellacasa and Gordon, 2007
Cephalocyclus luteolus (Horn, 1887)
Cephalocyclus majomaensis M. Dellacasa, G. Dellacasa and Gordon, 2013
Cephalocyclus mexicanus (Harold, 1862)
Cephalocyclus potosinus Dellacasa, Dellacasa and Gordon, 2007
Cephalocyclus pullatus (Schmidt, 1913)
Cephalocyclus rockefelleri Dellacasa, Dellacasa and Gordon, 2007
Chaunocolus cornutus Saylor, 1937
Chilothorax distinctus (Müller, 1776)
Chlorixanthe propinqua (Gory and Percheron, 1833)
Chnaunanthus chapini Saylor, 1937
Chnaunanthus discolor Burmeister, 1844
Chnaunanthus flavipennis (Horn, 1867)
Chrysina adelaida (Hope, 1840)
Chrysina beckeri Bates, 1889
Chrysina beyeri (Skinner, 1905)
Chrysina brevis (Rothschild and Jordan, 1894)
Chrysina erubescens Bates, 1889
Chrysina flohri (Ohaus, 1905)
Chrysina gloriosa (LeConte, 1854)
Chrysina lecontei (Horn, 1882)
Chrysina plusiotina (Ohaus, 1912)
Chrysina psittacina (Sturm, 1843)
Chrysina woodii (Horn, 1885)
Cinacanthus coquilletti (Linell, 1896)
Cinacanthus crenicollis (Fall, 1932)
Cinacanthus crescentus Gordon and Skelley, 2007
Cinacanthus hirsutus (Brown, 1928)
Cinacanthus militaris (LeConte, 1858)
Cinacanthus ulkei (Gordon, 2006)
Coelotrachelus kuntzeni (Schmidt, 1913)
Coelotrachelus rudis (LeConte, 1878)
Coelotrachelus symbius Gordon and Howden, 1973
Coenonycha acuta Cazier, 1943
Coenonycha ampla Cazier, 1943
Coenonycha barri Cazier, 1943
Coenonycha bowlesi Cazier, 1943
Coenonycha clementina Casey, 1909
Coenonycha clypeata McClay, 1943
Coenonycha crispata McClay, 1943
Coenonycha dimorpha Evans, 1986
Coenonycha fuga Cazier, 1943
Coenonycha fulva McClay, 1943
Coenonycha fusca McClay, 1943
Coenonycha globosa McClay, 1943
Coenonycha hageni Cazier, 1943
Coenonycha lurida Cazier, 1943
Coenonycha mediata Cazier, 1943
Coenonycha ochreata Evans, 1986
Coenonycha ovatis McClay, 1943
Coenonycha ovipennis Horn, 1876
Coenonycha pallida Cazier, 1943
Coenonycha parvula Fall, 1901
Coenonycha pascuensis Potts, 1945
Coenonycha purshiae Cazier, 1943
Coenonycha pygmaea Smith, 1986
Coenonycha rotundata (LeConte, 1856)
Coenonycha rubida McClay, 1943
Coenonycha santacruzae Evans, 1986
Coenonycha saylori Cazier, 1943
Coenonycha scotti McClay, 1943
Coenonycha sleeperi Evans, 1988
Coenonycha socialis Horn, 1876
Coenonycha stohleri Saylor, 1935
Coenonycha testacea Cazier, 1937
Coenonycha tingi Cazier, 1937
Coenonycha utahensis McClay, 1943
Colobopterus erraticus (Linnaeus, 1758)
Copris arizonensis Schaeffer, 1906
Copris fricator (Fabricius, 1787)
Copris gopheri Hubbard, 1894
Copris howdeni Matthews and Halffter, 1959
Copris incertus Say, 1835
Copris inemarginatus Blatchley, 1918
Copris klugi Harold, 1869
Copris laeviceps Harold, 1862
Copris lecontei Matthews, 1962
Copris lugubris Boheman, 1858
Copris macclevei Warner, 1990
Copris martinezi Matthews and Halffter, 1968
Copris minutus (Drury, 1773)
Copris moechus LeConte, 1854
Copris remotus LeConte, 1866
Coprophanaeus pluto (Harold, 1863)
Coscinocephalus cribrifrons (Schaeffer, 1906)
Cotalpa ashleyae LaRue, 1986
Cotalpa conclamara Young, 2002
Cotalpa consobrina Horn, 1871
Cotalpa flavida Horn, 1878
Cotalpa lanigera (Linnaeus, 1758)
Cotalpa subcribrata Wickham, 1905
Cotinis aliena Woodruff, 2008
Cotinis antonii (Dugés, 1878)
Cotinis boylei Goodrich, 1966
Cotinis impia (Fall, 1905)
Cotinis laticornis Bates, 1889
Cotinis mutabilis (Gory and Percheron, 1833)
Cotinis nitida (Linnaeus, 1758)
Cotinis orientalis Deloya and Ratcliffe, 1988
Cotinis producta Bates, 1889
Cotinis rufipennis Bates, 1889
Cotinis sinitoc Deloya, Ibañez-Bernal and Nogueira, 2000
Cremastocheilus academicus Krikken, 1982
Cremastocheilus angularis LeConte, 1857
Cremastocheilus armatus Walker, 1866
Cremastocheilus beameri Cazier, 1940
Cremastocheilus canaliculatus Kirby, 1827
Cremastocheilus castaneae Knoch, 1801
Cremastocheilus chapini Cazier, 1940
Cremastocheilus congener Casey, 1915
Cremastocheilus constricollis Cazier, 1940
Cremastocheilus crinitus LeConte, 1874
Cremastocheilus excavatus Cazier, 1940
Cremastocheilus harrisii Kirby, 1827
Cremastocheilus hirsutus Van Dyke, 1918
Cremastocheilus knochii LeConte, 1853
Cremastocheilus lengi Cazier, 1940
Cremastocheilus mentalis Cazier, 1940
Cremastocheilus mexicanus Schaum, 1841
Cremastocheilus nitens LeConte, 1853
Cremastocheilus opaculus Horn, 1894
Cremastocheilus planatus LeConte, 1863
Cremastocheilus planipes Horn, 1885
Cremastocheilus pulverulentus Cazier, 1940
Cremastocheilus puncticollis Cazier, 1940
Cremastocheilus quadratus Fall, 1912
Cremastocheilus quadricollis (Casey, 1915)
Cremastocheilus retractus LeConte, 1874
Cremastocheilus robinsoni Cazier, 1940
Cremastocheilus saucius LeConte, 1858
Cremastocheilus schaumii LeConte, 1853
Cremastocheilus setosifrons (Casey, 1915)
Cremastocheilus spinifer Horn, 1885
Cremastocheilus squamulosus LeConte, 1858
Cremastocheilus stathamae Cazier, 1961
Cremastocheilus tomentosus Warner, 1985
Cremastocheilus variolosus Kirby, 1826
Cremastocheilus westwoodi Horn, 1880
Cremastocheilus wheeleri LeConte, 1876
Cryptoscatomaseter acuminatus (Cartwright, 1944)
Cryptoscatomaseter brevicollis (LeConte, 1878)
Cryptoscatomaseter browni (Hinton, 1934)
Cryptoscatomaseter caccabatus (Warner and Skelley, 2006)
Cryptoscatomaseter captivus (Cartwright, 1944)
Cryptoscatomaseter criddlei (Brown, 1928)
Cryptoscatomaseter depressiusculus (Schmidt, 1907)
Cryptoscatomaseter durangoi (Gordon and Howden, 1973)
Cryptoscatomaseter ejectus Gordon and Skelley, 2007
Cryptoscatomaseter epulus (Gordon and Howden, 1973)
Cryptoscatomaseter excavatus (Robinson, 1940)
Cryptoscatomaseter gambrinus (Skelley and Gordon, 2002)
Cryptoscatomaseter haldemani (Horn, 1887)
Cryptoscatomaseter hubbelli (Skelley and Woodruff, 1991)
Cryptoscatomaseter incomptus Gordon and Skelley, 2007
Cryptoscatomaseter iowensis (Wickham, 1913)
Cryptoscatomaseter magnificens (Robinson, 1940)
Cryptoscatomaseter ochreipennis (Horn, 1871)
Cryptoscatomaseter oklahomensis (Brown, 1928)
Cryptoscatomaseter paulseni (Skelley, 2006)
Cryptoscatomaseter pholetus (Skelley and Woodruff, 1991)
Cryptoscatomaseter platypleurus (Skelley and Woodruff, 1991)
Cryptoscatomaseter punctissimus (Brown, 1928)
Cryptoscatomaseter rossi (Cartwright, 1944)
Cryptoscatomaseter salsburyi (Gordon, 2006)
Cryptoscatomaseter talpoidesi (Brown, 1928)
Cryptoscatomaseter umbricollis (Fall, 1907)
Cryptoscatomaseter utopensis (Warner and Skelley, 2006)
Cryptoscatomaseter yavapai (Robinson, 1946)
Cyclocephala aravaipensis Ratcliffe, 1992
Cyclocephala arenosa Howden and Endrödi, 1966
Cyclocephala barrerai Martínez, 1969
Cyclocephala borealis Arrow, 1911
Cyclocephala caelestis Delgado and Ratcliffe, 1990
Cyclocephala comata Bates, 1888
Cyclocephala deceptor (Casey, 1915)
Cyclocephala forcipulata Howden and Endrödi, 1966
Cyclocephala freudei Endrödi, 1963
Cyclocephala hirta LeConte, 1861
Cyclocephala knobelae (Brown, 1934)
Cyclocephala longula LeConte, 1863
Cyclocephala lunulata Burmeister, 1847
Cyclocephala lurida Bland, 1863
Cyclocephala mafaffa Burmeister, 1847
Cyclocephala melanocephala (Fabricius, 1775)
Cyclocephala miamiensis Howden and Endrödi, 1966
Cyclocephala nigricollis Burmeister, 1847
Cyclocephala parallela (Casey, 1915)
Cyclocephala pasadenae (Casey, 1915)
Cyclocephala pilosicollis Saylor, 1936
Cyclocephala puberula LeConte, 1863
Cyclocephala regularis Casey, 1915
Cyclocephala sanguinicollis Burmeister, 1847
Cyclocephala seditiosa LeConte, 1863
Cyclocephala sinaloae Howden and Endrödi, 1966
Cyclocephala sororia Bates, 1888
Cyclocephala sparsa Arrow, 1902
Cyclocephala testacea Burmeister, 1847
Cyclocephala variabilis Burmeister, 1847
Cyclocephala wandae Hardy, 1974
Dellacasiellus carpinterius Gordon and Skelley, 2007
Dellacasiellus claudus (Fall, 1932)
Dellacasiellus concavus (Say, 1823)
Dellacasiellus fucosus (Schmidt, 1922)
Dellacasiellus glamisensis Gordon and Skelley, 2007
Dellacasiellus griffini Gordon and Skelley, 2007
Dellacasiellus humboldti Gordon and Skelley, 2007
Dellacasiellus kirni (Cartwright, 1944)
Dellacasiellus laevigatus (Haldeman, 1848)
Dellacasiellus milleri Gordon and Skelley, 2007
Dellacasiellus pseudofucosus Gordon and Skelley, 2007
Dellacasiellus ruficlarus (Fall, 1932)
Deltochilum gibbosum (Fabricius, 1775)
Deltochilum scabriusculum Bates, 1887
Deltochilum tumidum Howden, 1966
Dialytellus dialytoides (Fall, 1907)
Dialytellus tragicus (Schmidt, 1916)
Dialytes criddlei Brown, 1929
Dialytes striatulus (Say, 1825)
Dialytes truncatus (Melsheimer, 1845)
Dialytes ulkei Horn, 1875
Dialytodius carri (Brown, 1929)
Dialytodius decipiens (Horn, 1887)
Dialytodius olympicus Gordon and Skelley, 2007
Dialytodius rogersi (Hatch, 1971)
Diapterna dugesi (Bates, 1887)
Diapterna hamata (Say, 1824)
Diapterna hyperborea (LeConte, 1850)
Diapterna omissa (LeConte, 1850)
Diapterna pinguella (Brown, 1929)
Diapterna pinguis (Haldeman, 1848)
Diastictus hawaiiensis Rakovic, 1981
Dichelonyx albicollis Burmeister, 1855
Dichelonyx backii Kirby, 1837
Dichelonyx canadensis Horn, 1876
Dichelonyx clypeata Horn, 1876
Dichelonyx crumbi Hatch, 1971
Dichelonyx decolorata Fall, 1901
Dichelonyx diluta Fall, 1901
Dichelonyx elongatula (Schönherr, 1817)
Dichelonyx fuscula LeConte, 1856
Dichelonyx kirbyi Brown, 1946
Dichelonyx lateralis Fall, 1901
Dichelonyx muscula Fall, 1901
Dichelonyx nana Fall, 1901
Dichelonyx pallens LeConte, 1856
Dichelonyx picea Horn, 1894
Dichelonyx pusilla LeConte, 1856
Dichelonyx robusta Fall, 1901
Dichelonyx subvittata LeConte, 1856
Dichelonyx sulcata LeConte, 1856
Dichelonyx testaceipennis Fall, 1907
Dichelonyx truncata LeConte, 1856
Dichelonyx vaga Fall, 1901
Dichelonyx valida LeConte, 1856
Dichelonyx vandykei Saylor, 1932
Dichelonyx vicina Fall, 1901
Dichotomius carolinus (Linnaeus, 1767)
Dichotomius colonicus (Say, 1835)
Dinacoma caseyi Blaisdell, 1930
Dinacoma marginata (Casey, 1886)
Diplotaxis abnormis Fall, 1909
Diplotaxis academia Vaurie, 1960
Diplotaxis aenea Blanchard, 1851
Diplotaxis aequalis Cazier, 1940
Diplotaxis ambigua Fall, 1909
Diplotaxis angularis LeConte, 1856
Diplotaxis anthracina Fall, 1909
Diplotaxis anxia (LeConte, 1856)
Diplotaxis arctifrons Bates, 1888
Diplotaxis arizonica Schaeffer, 1907
Diplotaxis atlantis Fall, 1909
Diplotaxis atratula LeConte, 1856
Diplotaxis aulacochela Cazier, 1940
Diplotaxis australis Vaurie, 1960
Diplotaxis bakeri Cazier, 1940
Diplotaxis barbarae Vaurie and Cazier, 1955
Diplotaxis basalis Fall, 1909
Diplotaxis belfragei Fall, 1909
Diplotaxis beyeri Schaeffer, 1907
Diplotaxis bidentata LeConte, 1856
Diplotaxis blanchardi Vaurie, 1956
Diplotaxis boops Bates, 1888
Diplotaxis bowditchi Fall, 1909
Diplotaxis brachyptera Vaurie, 1960
Diplotaxis brevicollis LeConte, 1856
Diplotaxis brevicornis Cazier, 1940
Diplotaxis brevidens LeConte, 1856
Diplotaxis brevisetosa Linell, 1896
Diplotaxis caelestis Delgado and Capistran, 1993
Diplotaxis carbonata LeConte, 1856
Diplotaxis catarinas Vaurie and Cazier, 1955
Diplotaxis chiricahuae Fall, 1909
Diplotaxis circulans Vaurie, 1960
Diplotaxis coenonychoides Davidson and Davidson, 2006
Diplotaxis commixta Vaurie, 1960
Diplotaxis completa Cazier, 1940
Diplotaxis conformis Fall, 1909
Diplotaxis confusa Fall, 1909
Diplotaxis connata Schaeffer, 1905
Diplotaxis contracta Bates, 1888
Diplotaxis convexilabra Vaurie and Cazier, 1955
Diplotaxis corbula Vaurie, 1960
Diplotaxis coriacea Bates, 1888
Diplotaxis corvina LeConte, 1856
Diplotaxis costanera Vaurie, 1958
Diplotaxis cribatella Bates, 1889
Diplotaxis cribulosa LeConte, 1856
Diplotaxis curvaticeps Fall, 1909
Diplotaxis dahli Cazier, 1940
Diplotaxis decima Vaurie and Cazier, 1955
Diplotaxis dentella Fall, 1909
Diplotaxis denticeps Bates, 1887
Diplotaxis deserta Fall, 1932
Diplotaxis dissona Vaurie, 1960
Diplotaxis dubia LeConte, 1856
Diplotaxis durango Vaurie, 1960
Diplotaxis elongata Vaurie, 1960
Diplotaxis errans Fall, 1909
Diplotaxis exstans Vaurie, 1960
Diplotaxis fimbriata Fall, 1909
Diplotaxis fissilabris Fall, 1909
Diplotaxis fissilis Vaurie and Cazier, 1955
Diplotaxis flexa Vaurie, 1960
Diplotaxis fossipalpa Fall, 1909
Diplotaxis frondicola (Say, 1825)
Diplotaxis fulva (LeConte, 1856)
Diplotaxis glabrimargo Vaurie and Cazier, 1955
Diplotaxis harperi Blanchard, 1851
Diplotaxis haydenii LeConte, 1856
Diplotaxis hebes Bates, 1888
Diplotaxis hirsuta Vaurie, 1958
Diplotaxis illustris Fall, 1909
Diplotaxis impar Vaurie and Cazier, 1955
Diplotaxis incuria Fall, 1932
Diplotaxis indigena Vaurie and Cazier, 1955
Diplotaxis ingenua Fall, 1909
Diplotaxis insignis LeConte, 1861
Diplotaxis knausii Schaeffer, 1907
Diplotaxis kuschel Vaurie, 1960
Diplotaxis languida LeConte, 1878
Diplotaxis lengii Fall, 1909
Diplotaxis levicosta Fall, 1909
Diplotaxis liberta (Germar, 1824)
Diplotaxis macronycha Fall, 1909
Diplotaxis magna Bates, 1888
Diplotaxis marginicollis Fall, 1909
Diplotaxis martinezi Delgado and Capistran, 1993
Diplotaxis mascula Vaurie, 1960
Diplotaxis maura Fall, 1909
Diplotaxis megapleura Vaurie, 1958
Diplotaxis mentalis Fall, 1909
Diplotaxis mima Vaurie and Cazier, 1955
Diplotaxis mimosae Fall, 1909
Diplotaxis misella Fall, 1909
Diplotaxis missionaria Vaurie, 1960
Diplotaxis moerens LeConte, 1856
Diplotaxis muricata Schaeffer, 1907
Diplotaxis mus Fall, 1932
Diplotaxis nigriventris Bates, 1887
Diplotaxis obregon Vaurie, 1958
Diplotaxis obscura LeConte, 1859
Diplotaxis ohausi Moser, 1921
Diplotaxis pacata LeConte, 1856
Diplotaxis pala Vaurie, 1960
Diplotaxis parallela Fall, 1909
Diplotaxis parpolita Vaurie, 1960
Diplotaxis parvicollis Fall, 1909
Diplotaxis persisae Cazier, 1940
Diplotaxis pilipennis Moser, 1918
Diplotaxis pinalica Fall, 1932
Diplotaxis planidens Fall, 1909
Diplotaxis polita Fall, 1909
Diplotaxis profunda Vaurie and Cazier, 1955
Diplotaxis puberula LeConte, 1863
Diplotaxis pubipes Schaeffer, 1907
Diplotaxis pumila Fall, 1909
Diplotaxis punctata LeConte, 1856
Diplotaxis punctatorugosa Blanchard, 1851
Diplotaxis puncticeps Moser, 1918
Diplotaxis punctipennis LeConte, 1856
Diplotaxis punctulata Horn, 1894
Diplotaxis pygidialis Vaurie, 1960
Diplotaxis residua Fall, 1909
Diplotaxis rex Vaurie, 1958
Diplotaxis robertmarki Davidson and Davidson, 2006
Diplotaxis rockefelleri Vaurie and Cazier, 1955
Diplotaxis rosae Vaurie and Cazier, 1955
Diplotaxis rotunda Vaurie, 1960
Diplotaxis rudis (LeConte, 1859)
Diplotaxis rufa Linell, 1895
Diplotaxis rufiola Fall, 1909
Diplotaxis saltensis Vaurie, 1960
Diplotaxis saylori Cazier, 1940
Diplotaxis schaefferi Fall, 1909
Diplotaxis semifoveata Fall, 1909
Diplotaxis sierrae Fall, 1909
Diplotaxis simplex Blanchard, 1851
Diplotaxis sonora Vaurie, 1960
Diplotaxis sordida (Say, 1825)
Diplotaxis sparsa Vaurie, 1960
Diplotaxis statura Cazier, 1940
Diplotaxis subangulata LeConte, 1856
Diplotaxis subcostata Blanchard, 1851
Diplotaxis subrugata Moser, 1918
Diplotaxis sulcatula Fall, 1909
Diplotaxis superflua Vaurie, 1960
Diplotaxis tarsalis Schaeffer, 1907
Diplotaxis tenebrosa Fall, 1909
Diplotaxis tenuis LeConte, 1856
Diplotaxis texana LeConte, 1856
Diplotaxis thoracica Fall, 1909
Diplotaxis trapezifera Bates, 1887
Diplotaxis trementina Fall, 1909
Diplotaxis tristis Kirby, 1837
Diplotaxis truncatula LeConte, 1856
Diplotaxis ungulata Cazier, 1940
Diplotaxis urbana Vaurie, 1960
Diplotaxis vandykei Vaurie, 1958
Drepanacanthoides larreae (Horn, 1887)
Drepanacanthoides walshii (Horn, 1870)
Dynastes granti Horn, 1870
Dynastes hyllus Chevrolat, 1843
Dynastes tityus (Linnaeus, 1767)
Dyscinetus laevicollis Arrow, 1937
Dyscinetus morator (Fabricius, 1798)
Dyscinetus picipes (Burmeister, 1847)
Ectinoplectron oryctoides (Ohaus, 1905)
Enema endymion Chevrolat, 1843
Euetheola humilis (Burmeister, 1847)
Euetheola subglabra (Schaeffer, 1909)
Euoniticellus africanus (Harold, 1873)
Euoniticellus cubiensis (Laporte, 1840)
Euoniticellus intermedius (Reiche, 1849)
Euparia castanea LePeletier and Serville, 1828
Euparixia duncani Brown, 1927
Euparixia mexicana Gordon and McCleve, 2003
Euparixia moseri Woodruff and Cartwright, 1967
Euphoria anneae (Howden, 1955)
Euphoria areata (Fabricius, 1775)
Euphoria basalis (Gory and Percheron, 1833)
Euphoria biguttata (Gory and Percheron, 1833)
Euphoria bispinis (Bates, 1889)
Euphoria canescens (Gory and Percheron, 1833)
Euphoria casselberryi Robinson, 1937
Euphoria devulsa Horn, 1880
Euphoria discicollis (Thomson, 1878)
Euphoria fascifera (LeConte, 1861)
Euphoria fulgida (Fabricius, 1775)
Euphoria geminata (Chevrolat, 1835)
Euphoria herbacea (Olivier, 1789)
Euphoria hirtipes Horn, 1880
Euphoria histrionica Thomson, 1878
Euphoria inda (Linnaeus, 1758)
Euphoria kernii Haldeman, 1852
Euphoria leucopyge Bates, 1889
Euphoria lineoligera (Blanchard, 1850)
Euphoria monticola Bates, 1889
Euphoria pilipennis (Kraatz, 1883)
Euphoria pulchella (Gory and Percheron, 1833)
Euphoria quadricollis Bates, 1889
Euphoria schottii (LeConte, 1854)
Euphoria sepulchralis (Fabricius, 1801)
Euphoria verticalis Horn, 1880
Eupleurus subterraneus (Linnaeus, 1758)
Flaviellus aggregatus (Gordon, 1977)
Flaviellus gordoni (Ratcliffe, 1988)
Flaviellus perfimbriatus (Gordon, 1977)
Flaviellus phalerioides (Horn, 1870)
Flaviellus phalerius Gordon and Skelley, 2007
Flaviellus subtruncatus (LeConte, 1878)
Fossocarus creoleorum Howden, 1961
Genuchinus ineptus (Horn, 1885)
Geomyphilus alabama (Skelley and Gordon, 1995)
Geomyphilus essigi (Saylor, 1935)
Geomyphilus geronimo Gordon and Skelley, 2007
Geomyphilus insolitus (Brown, 1928)
Geomyphilus kiowensis (Gordon and Salsbury, 1999)
Geomyphilus leptotarsis (Brown, 1928)
Geomyphilus pinolus Gordon and Skelley, 2007
Geomyphilus rubiginosus (Horn, 1870)
Geomyphilus russeus (Brown, 1928)
Geomyphilus thomomysi (Brown, 1928)
Geomyphilus ungulatus (Fall, 1901)
Geomyphilus viceversus (Paulsen, 2006)
Geopsammodius fuscus Skelley, 2006
Geopsammodius hydropicus (Horn, 1887)
Geopsammodius morrisi Skelley, 2006
Geopsammodius ohoopee Skelley, 2006
Geopsammodius relictillus (Deyrup and Woodruff, 1991)
Geopsammodius rileyi Skelley, 2006
Geopsammodius subpedalis Skelley, 2006
Geopsammodius unsidensis Skelley, 2006
Geopsammodius withlacoochee Skelley, 2006
Gillaspytes janzeni Howden, 1980
Gnorimella maculosa (Knoch, 1801)
Golofa imperialis Thomson, 1858
Golofa xiximeca Morón, 1995
Gronocarus autumnalis Schaeffer, 1927
Gronocarus inornatus Skelley, 2005
Gymnetina cretacea (LeConte, 1866)
Gymnetis caseyi Antoine, 2001
Gymnopyge coquilletti Linell, 1896
Gymnopyge hirsuta Cazier, 1939
Gymnopyge hopliaeformis Linell, 1896
Gymnopyge pygmaea Linell, 1896
Halffterinetis gonzaloi Morón and Nogueira, 2007
Halffterinetis violetae Morón and Nogueira, 2007
Haroldiataenius buvexus Stebnicka and Skelley, 2009
Haroldiataenius convexus (Robinson, 1940)
Haroldiataenius griffini (Cartwright, 1974)
Haroldiataenius hintoni (Saylor, 1933)
Haroldiataenius limbatus (Bates, 1887)
Haroldiataenius lucanus (Horn, 1871)
Haroldiataenius mariarum (Bates, 1887)
Haroldiataenius saramari (Cartwright, 1939)
Haroldiataenius semipilosus (Van Dyke, 1928)
Haroldiellus lansbergei (Harold, 1874)
Haroldiellus sallei (Harold, 1863)
Hemiphileurus illatus (LeConte, 1854)
Hologymnetis argenteola (Bates, 1889)
Hologymnetis cinerea (Gory and Percheron, 1833)
Homoiosternus beckeri Ohaus, 1901
Hoplia callipyge LeConte, 1856
Hoplia cinereonebulosa Nonfried, 1895
Hoplia dispar LeConte, 1880
Hoplia equina LeConte, 1880
Hoplia floridana Fisher, 1918
Hoplia gilleti Hardy, 1977
Hoplia hirta LeConte, 1880
Hoplia laticollis LeConte, 1856
Hoplia modesta Haldeman, 1843
Hoplia mucorea (Germar, 1824)
Hoplia sackenii LeConte, 1880
Hoplia trifasciata Say, 1825
Hoplia trivialis Harold, 1869
Hornietus ventralis (Horn, 1887)
Hypothyce burnei Skelley, 2005
Hypothyce mixta Howden, 1968
Hypothyce osburni (Cartwright, 1967)
Hypotrichia spissipes LeConte, 1861
Inca clathrata (Olivier, 1792)
Irrasinus dumasi Gordon and Skelley, 2007
Irrasinus stupidus (Horn, 1870)
Isonychus arizonensis Howden, 1959
Labarrus lividus (Olivier, 1789)
Labarrus pseudolividus Balthasar, 1941
Lechorodius lutulentus (Haldeman, 1843)
Lechorodius terminalis (Say, 1823)
Leiopsammodius acei Harpootlian, Gordon and Woodruff, 2000
Leiopsammodius deyrupi Harpootlian, Gordon and Woodruff, 2000
Leiopsammodius malkini (Cartwright, 1946)
Leiopsammodius ocmulgeei Harpootlian, Gordon and Woodruff, 2000
Leptohoplia testaceipennis Saylor, 1935
Liatongus californicus (Horn, 1882)
Lichnia gallardoi Gutiérrez, 1943
Lichnia limbata Erichson, 1835
Liothorax alternatus (Horn, 1870)
Liothorax consociatus (Horn, 1887)
Liothorax innexus (Say, 1835)
Liothorax subaeneus (LeConte, 1857)
Lissomelas flohri Bates, 1889
Luxolinus luxatus (Horn, 1887)
Macrodactylus angustatus (Palisot de Beauvois, 1805)
Macrodactylus fulvescens Bates, 1887
Macrodactylus impressus Bates, 1887
Macrodactylus infuscatus Bates, 1887
Macrodactylus lineaticollis Bates, 1887
Macrodactylus mexicanus Burmeister, 1855
Macrodactylus murinus Bates, 1887
Macrodactylus nigripes Bates, 1887
Macrodactylus rufescens Bates, 1887
Macrodactylus silaonus Bates, 1887
Macrodactylus subspinosus (Fabricius, 1775)
Macrodactylus uniformis Horn, 1876
Macrodactylus variipes Bates, 1887
Macrodactylus virens Bates, 1887
Maculaphodius conspersus (Horn, 1887)
Maladera castanea (Arrow, 1913)
Malagoniella astyanax (Olivier, 1789)
Martineziana dutertrei (Chalumeau, 1983)
Martineziana vandykei (Hinton, 1936)
Megasoma cedrosum Hardy, 1972
Megasoma lecontei Hardy, 1972
Megasoma lenczyi Cartwright, 1976
Megasoma occidentalis Bolívar y Pieltain, Jiménez-Asúa and Martínez, 1963
Megasoma pachecoi Cartwright, 1963
Megasoma punctulatum Cartwright, 1952
Megasoma sleeperi Hardy, 1972
Megasoma thersites LeConte, 1861
Megasoma vogti Cartwright, 1963
Melanocanthon bispinatus (Robinson, 1941)
Melanocanthon granulifer (Schmidt, 1920)
Melanocanthon nigricornis (Say, 1823)
Melanocanthon punctaticollis (Schaeffer, 1915)
Melinopterus consentaneus (LeConte, 1850)
Melinopterus femoralis (Say, 1823)
Melinopterus prodromus (Brahm, 1790)
Mendidius aculeatus (Robinson, 1940)
Merogyrus rotundiceps (Fall, 1907)
Micraegialia pusilla (Horn, 1887)
Nefoncerus convergens (Horn, 1894)
Neopsammodius blandus (Fall, 1932)
Neopsammodius interruptus (Say, 1835)
Neopsammodius mimeticus (Fall, 1932)
Neopsammodius quinqueplicatus (Horn, 1871)
Neopsammodius saltilloensis (Cartwright, 1955)
Neopsammodius werneri (Cartwright, 1955)
Neotrichonotulus inurbanus (Gordon and Howden, 1973)
Nialaphodius nigritus (Fabricius, 1801)
Nipponoserica peregrina (Chapin, 1938)
Odontolytes denominatus (Chevrolat, 1864)
Odontopsammodius armaticeps (Fall, 1932)
Odontopsammodius cruentus (Harold, 1867)
Oncerus floralis LeConte, 1856
Oniticellus cinctus (Fabricius, 1775)
Oniticellus militaris (Castelnau, 1840)
Oniticellus rhinocerulus Bates, 1889
Onitis alexis Klug, 1835
Onitis vanderkelleni Lansberge, 1886
Ontherus mexicanus Harold, 1868
Onthophagus aciculatulus Blatchley, 1928
Onthophagus alluvius Howden and Cartwright, 1963
Onthophagus antivagans Howden and Génier, 2004
Onthophagus arnetti Howden and Cartwright, 1963
Onthophagus batesi Howden and Cartwright, 1963
Onthophagus binodis (Thunberg, 1818)
Onthophagus brevifrons Horn, 1881
Onthophagus browni Howden and Cartwright, 1963
Onthophagus canelasensis Howden and Génier, 2004
Onthophagus cartwrighti Howden, 1973
Onthophagus catta (Fabricius, 1787)
Onthophagus cavernicollis Howden and Cartwright, 1963
Onthophagus chevrolati Harold, 1869
Onthophagus coahuilae Zunino and Halffter, 1988
Onthophagus cochisus Brown, 1927
Onthophagus concinnus Laporte, 1840
Onthophagus coproides Horn, 1881
Onthophagus cribricollis Horn, 1881
Onthophagus cuevensis Howden, 1973
Onthophagus cyanellus Bates, 1887
Onthophagus cynomysi Brown, 1927
Onthophagus depressus Harold, 1871
Onthophagus foliaceus Lansberge, 1886
Onthophagus fuscus Boucomont, 1932
Onthophagus gazella (Fabricius, 1787)
Onthophagus hecate (Panzer, 1794)
Onthophagus hidalgus Zunino and Halffter, 1988
Onthophagus hoepfneri Harold, 1869
Onthophagus incensus Say, 1835
Onthophagus knausi Brown, 1927
Onthophagus knulli Howden and Cartwright, 1963
Onthophagus landolti Harold, 1880
Onthophagus lecontei Harold, 1871
Onthophagus mcclevei Howden and Génier, 2004
Onthophagus medorensis Brown, 1929
Onthophagus mextexus Howden and Cartwright, 1970
Onthophagus nigriventris d'Orbigny, 1905
Onthophagus nuchicornis (Linnaeus, 1758)
Onthophagus oklahomensis Brown, 1927
Onthophagus orpheus (Panzer, 1794)
Onthophagus pennsylvanicus Harold, 1871
Onthophagus polyphemi Hubbard, 1894
Onthophagus pseudofuscus Zunino and Halffter, 1988
Onthophagus saggitarius (Fabricius, 1775)
Onthophagus schaefferi Howden and Cartwright, 1963
Onthophagus striatulus (Palisot de Beauvois, 1809)
Onthophagus subaeneus (Palisot de Beauvois, 1811)
Onthophagus subopacus Robinson, 1940
Onthophagus subtropicus Howden and Cartwright, 1963
Onthophagus taurus (Schreber, 1759)
Onthophagus tuberculifrons Harold, 1871
Onthophagus velutinus Horn, 1875
Orizabus batesi Prell, 1914
Orizabus brevicollis Prell, 1914
Orizabus clunalis (LeConte, 1856)
Orizabus isodonoides Fairmaire, 1878
Orizabus ligyroides Horn, 1885
Orizabus mcclevei Warner, 2011
Orizabus pinalicus Warner, 2011
Orizabus pyriformis (LeConte, 1847)
Orizabus ratcliffei Delgado, 2008
Orizabus rubricollis Prell, 1914
Oscarinus abusus (Fall, 1907)
Oscarinus bottimeri (Cartwright, 1957)
Oscarinus brimleyi (Cartwright, 1939)
Oscarinus cabreroi M. Dellacasa, G. Dellacasa and Gordon, 2013
Oscarinus crassuloides (Fall, 1907)
Oscarinus crassulus (Horn, 1870)
Oscarinus floridanus (Robinson, 1947)
Oscarinus lodingi (Cartwright, 1957)
Oscarinus matiganae (Paulsen, 2006)
Oscarinus odocoilis (Robinson, 1939)
Oscarinus pseudabusus (Cartwright, 1957)
Oscarinus rusicola (Melsheimer, 1845)
Oscarinus silvanicus (Cartwright, 1972)
Oscarinus stuessyi Gordon and Skelley, 2007
Oscarinus texensis (Cartwright, 1972)
Oscarinus welderi Gordon and Skelley, 2007
Oscarinus windsori (Cartwright, 1939)
Osmoderma eremicola (Knoch, 1801)
Osmoderma scabra (Palisot de Beauvois, 1805)
Osmoderma subplanata Casey, 1915
Otophorus haemorrhoidalis (Linnaeus, 1758)
Oxygrylius peninsularis Casey, 1915
Oxygrylius ruginasus (LeConte, 1856)
Oxyomus silvestris (Scopoli, 1763)
Parabyrsopolis chihuahuae (Bates, 1888)
Parachrysina borealis Jameson, 1991
Paracotalpa deserta Saylor, 1940
Paracotalpa granicollis (Haldeman, 1852)
Paracotalpa puncticollis (LeConte, 1863)
Paracotalpa ursina (Horn, 1867)
Parapsammodius bidens (Horn, 1871)
Parapsammodius puncticollis (LeConte, 1858)
Parastasia brevipes (LeConte, 1856)
Parataenius simulator (Harold, 1868)
Pardalosus neodistinctus (Brown, 1928)
Pardalosus pardalis (LeConte, 1857)
Pardalosus pseudopardalis Gordon and Skelley, 2007
Pardalosus pumilio (Schmidt, 1907)
Pardalosus sayi (Gordon, 2006)
Pardalosus serval (Say, 1835)
Pardalosus slevini (Van Dyke, 1928)
Pelidnota lucae LeConte, 1863
Pelidnota lugubris LeConte, 1874
Pelidnota perplexa Hardy, 1975
Pelidnota punctata (Linnaeus, 1758)
Pelidnota strigosa Laporte, 1840
Pelidnota virescens Burmeister, 1844
Phaeaphodius rectus (Motschulsky, 1866)
Phanaeus adonis Harold, 1863
Phanaeus amethystinus Harold, 1863
Phanaeus amithaon Harold, 1875
Phanaeus daphnis Harold, 1863
Phanaeus difformis LeConte, 1847
Phanaeus flohri Nevinson, 1892
Phanaeus furiosus Bates, 1887
Phanaeus genieri Arnaud, 2001
Phanaeus igneus MacLeay, 1819
Phanaeus palliatus Sturm, 1843
Phanaeus quadridens (Say, 1835)
Phanaeus triangularis (Say, 1823)
Phanaeus vindex MacLeay, 1819
Phanaeus yecoraensis Edmonds, 2004
Phileurus truncatus (Palisot de Beauvois, 1806)
Phileurus valgus (Olivier, 1789)
Phobetus chearyi Hardy, 1973
Phobetus ciliatus Barrett, 1935
Phobetus comatus LeConte, 1856
Phobetus desertus Blom and Clark, 1984
Phobetus humeralis Cazier, 1937
Phobetus mojavus Barrett, 1933
Phobetus palpalis Saylor, 1936
Phobetus panamintensis Hardy, 1978
Phobetus saylori Cazier, 1937
Phobetus sleeperi Hardy, 1978
Phobetus testaceus LeConte, 1861
Phyllophaga aemula (Horn, 1887)
Phyllophaga aequalis (LeConte, 1854)
Phyllophaga affabilis (Horn, 1887)
Phyllophaga albina (Burmeister, 1855)
Phyllophaga almada Saylor, 1941
Phyllophaga anomaloides (Bates, 1888)
Phyllophaga antennata (Smith, 1889)
Phyllophaga anxia (LeConte, 1850)
Phyllophaga apicata Reinhard, 1939
Phyllophaga aragoni Morón, 2013
Phyllophaga arcta (Horn, 1887)
Phyllophaga arenicola Howden, 1960
Phyllophaga arizona Saylor, 1940
Phyllophaga arkansana (Schaeffer, 1906)
Phyllophaga aurea Luginbill and Painter, 1941
Phyllophaga babicora Morón, 2004
Phyllophaga balia (Say, 1825)
Phyllophaga barda (Horn, 1887)
Phyllophaga beckeri (Moser, 1921)
Phyllophaga bellamyi Warner and Morón, 1992
Phyllophaga bilobatata Saylor, 1939
Phyllophaga bipartita (Horn, 1887)
Phyllophaga blanchardi (Arrow, 1933)
Phyllophaga blanda Sanderson, 1958
Phyllophaga bottimeri Reinhard, 1950
Phyllophaga brevidens (Bates, 1888)
Phyllophaga bruneri Chapin, 1932
Phyllophaga buapae Morón, 2013
Phyllophaga bueta Saylor, 1940
Phyllophaga cahitana Morón, 2001
Phyllophaga calceata (LeConte, 1856)
Phyllophaga calculiventris Saylor, 1935
Phyllophaga callosiventris (Moser, 1921)
Phyllophaga canipolea Saylor, 1948
Phyllophaga carminator (Horn, 1894)
Phyllophaga cavata (Bates, 1888)
Phyllophaga cazahuata Morón, 2013
Phyllophaga cazieriana Saylor, 1938
Phyllophaga certanca Saylor, 1943
Phyllophaga chapini Saylor, 1940
Phyllophaga chippewa Saylor, 1939
Phyllophaga ciudadensis (Bates, 1888)
Phyllophaga clemens (Horn, 1887)
Phyllophaga clypeata (Horn, 1887)
Phyllophaga cochisa Saylor, 1940
Phyllophaga congrua (LeConte, 1856)
Phyllophaga contaminata Fall, 1932
Phyllophaga corrosa (LeConte, 1856)
Phyllophaga crassissima (Blanchard, 1851)
Phyllophaga crenulata (Frölich, 1792)
Phyllophaga cribrosa (LeConte, 1853)
Phyllophaga crinita (Burmeister, 1855)
Phyllophaga cristagalli (Arrow, 1933)
Phyllophaga cupuliformis Langston, 1924
Phyllophaga curialis Reinhard, 1939
Phyllophaga cushmani Saylor, 1940
Phyllophaga cylindrica (Burmeister, 1855)
Phyllophaga davisi Langston, 1927
Phyllophaga debilis (LeConte, 1856)
Phyllophaga delata (Horn, 1887)
Phyllophaga densicollis (LeConte, 1863)
Phyllophaga dentex (Bates, 1888)
Phyllophaga diffinis (Blanchard, 1851)
Phyllophaga dispar (Burmeister, 1855)
Phyllophaga disparilis (Horn, 1878)
Phyllophaga divertens (Bates, 1888)
Phyllophaga drakii (Kirby, 1837)
Phyllophaga duncani (Barrett, 1933)
Phyllophaga durango Saylor, 1940
Phyllophaga durangoana (Moser, 1921)
Phyllophaga durangosa Saylor, 1943
Phyllophaga ecostata (Horn, 1887)
Phyllophaga eligia Sanderson, 1958
Phyllophaga elizoria Saylor, 1937
Phyllophaga elongata (Linell, 1896)
Phyllophaga eniba Saylor, 1943
Phyllophaga ephilida (Say, 1825)
Phyllophaga epigaea (Wickham, 1903)
Phyllophaga errans (LeConte, 1860)
Phyllophaga estacea Saylor, 1943
Phyllophaga falsa (LeConte, 1856)
Phyllophaga falta (LeConte, 1856)
Phyllophaga farcta (LeConte, 1856)
Phyllophaga fervida (Fabricius, 1775)
Phyllophaga fimbripes (LeConte, 1856)
Phyllophaga flavipennis (Horn, 1885)
Phyllophaga floridana Robinson, 1938
Phyllophaga foralita Saylor, 1938
Phyllophaga forbesi Glasgow, 1916
Phyllophaga forsteri (Burmeister, 1855)
Phyllophaga foxii Davis, 1920
Phyllophaga fraterna Harris, 1842
Phyllophaga fucata (Horn, 1887)
Phyllophaga fulviventris (Moser, 1918)
Phyllophaga fusca (Frölich, 1792)
Phyllophaga futilis (LeConte, 1850)
Phyllophaga gaigei Sanderson, 1948
Phyllophaga galeanae Saylor, 1943
Phyllophaga gentryi (Saylor, 1936)
Phyllophaga georgiana (Horn, 1885)
Phyllophaga ghentata (Saylor, 1937)
Phyllophaga gigantissima Saylor, 1935
Phyllophaga glaberrima (Blanchard, 1851)
Phyllophaga glabricula (LeConte, 1856)
Phyllophaga gracilis (Burmeister, 1855)
Phyllophaga granti Saylor, 1940
Phyllophaga gravidula (Moser, 1921)
Phyllophaga hamata (Horn, 1887)
Phyllophaga hirsuta (Knoch, 1801)
Phyllophaga hirticula (Knoch, 1801)
Phyllophaga hirtiventris (Horn, 1887)
Phyllophaga hornii (Smith, 1889)
Phyllophaga huachuca Saylor, 1940
Phyllophaga hubbelli Cartwright, 1946
Phyllophaga idonea Sanderson, 1948
Phyllophaga ignava (Horn, 1887)
Phyllophaga ilicis (Knoch, 1801)
Phyllophaga impar Davis, 1920
Phyllophaga implicita (Horn, 1887)
Phyllophaga incuria Sanderson, 1942
Phyllophaga inepta (Horn, 1887)
Phyllophaga infidelis (Horn, 1887)
Phyllophaga inflexa Barrett, 1935
Phyllophaga instabilis Blackwelder, 1944
Phyllophaga inversa (Horn, 1887)
Phyllophaga invisa Riley and Wolfe, 1997
Phyllophaga juvenilis (Fall, 1932)
Phyllophaga karlsioei (Linell, 1898)
Phyllophaga kentuckiana Ritcher, 1937
Phyllophaga knausii (Schaeffer, 1907)
Phyllophaga knochii (Schönherr and Gyllenhal, 1817)
Phyllophaga koehleriana Saylor, 1940
Phyllophaga lalanza Saylor, 1941
Phyllophaga lanceolata (Say, 1824)
Phyllophaga laportaei (Blanchard, 1851)
Phyllophaga latidens (Schaeffer, 1906)
Phyllophaga latifrons (LeConte, 1856)
Phyllophaga lenis (Horn, 1887)
Phyllophaga lobata (Fall, 1908)
Phyllophaga lodingi Sanderson, 1939
Phyllophaga longispina (Smith, 1889)
Phyllophaga longitarsa (Say, 1824)
Phyllophaga lota Luginbill, 1928
Phyllophaga luctuosa (Horn, 1887)
Phyllophaga luginbilli Saylor, 1941
Phyllophaga macgregori Morón, 2004
Phyllophaga macmurryi Saylor, 1940
Phyllophaga maculicollis (LeConte, 1863)
Phyllophaga magnicornis (Moser, 1921)
Phyllophaga marginalis (LeConte, 1856)
Phyllophaga mariana Fall, 1929
Phyllophaga meadei Saylor, 1940
Phyllophaga micans (Knoch, 1801)
Phyllophaga michelbacheri Saylor, 1940
Phyllophaga microcera (Arrow, 1933)
Phyllophaga micros (Bates, 1888)
Phyllophaga mimicana Saylor, 1938
Phyllophaga miraflora Saylor, 1940
Phyllophaga misteca (Bates, 1888)
Phyllophaga molopia (Bates, 1888)
Phyllophaga monstrosa (Saylor, 1935)
Phyllophaga mucorea (LeConte, 1856)
Phyllophaga nebulosa Polihronakis, 2007
Phyllophaga neomexicana Saylor, 1940
Phyllophaga nitida (LeConte, 1856)
Phyllophaga nogales Saylor, 1940
Phyllophaga nogueirana Morón, 2002
Phyllophaga nosa Blackwelder, 1944
Phyllophaga obsoleta (Blanchard, 1851)
Phyllophaga okeechobea Robinson, 1948
Phyllophaga omani Sanderson, 1937
Phyllophaga opaca (Moser, 1918)
Phyllophaga opacicollis (Horn, 1878)
Phyllophaga ovalis (Cartwright, 1939)
Phyllophaga pallida (Horn, 1885)
Phyllophaga panorpa Sanderson, 1950
Phyllophaga parilis (Bates, 1888)
Phyllophaga parvidens (LeConte, 1856)
Phyllophaga paternoi Glasgow, 1925
Phyllophaga pearliae Davis, 1920
Phyllophaga peninsularis Saylor, 1940
Phyllophaga pentaphylla (Bates, 1888)
Phyllophaga perlonga Davis, 1920
Phyllophaga pilosipes Saylor, 1940
Phyllophaga pilositarsis Blackwelder, 1944
Phyllophaga planeta Reinhard, 1950
Phyllophaga plena (Fall, 1932)
Phyllophaga pleroma Reinhard, 1940
Phyllophaga postrema (Horn, 1887)
Phyllophaga praesidii (Bates, 1888)
Phyllophaga praetermissa (Horn, 1887)
Phyllophaga profunda (Blanchard, 1851)
Phyllophaga pruinosa (Blanchard, 1851)
Phyllophaga prunina (LeConte, 1856)
Phyllophaga prununculina (Burmeister, 1855)
Phyllophaga pseudofloridana Woodruff and Beck, 1989
Phyllophaga psiloptera Sanderson, 1939
Phyllophaga pudorosa Reinhard, 1939
Phyllophaga pulcher (Linell, 1896)
Phyllophaga pusillidens Fall, 1937
Phyllophaga querca (Knoch, 1801)
Phyllophaga quetzala Morón, 2001
Phyllophaga ravida (Blanchard, 1851)
Phyllophaga reevesi Saylor, 1939
Phyllophaga regiomontana Morón, 2001
Phyllophaga reinhardi Saylor, 1940
Phyllophaga renodis Reinhard, 1939
Phyllophaga riviera Reinhard, 1950
Phyllophaga rolbakeri Saylor, 1940
Phyllophaga rolstoni Riley and Wolfe, 1997
Phyllophaga rossi Saylor, 1939
Phyllophaga rubiginosa (LeConte, 1856)
Phyllophaga rubricosa Reinhard, 1939
Phyllophaga rugipennis (Schaufuss, 1858)
Phyllophaga rugosa (Melsheimer, 1845)
Phyllophaga sacoma Reinhard, 1939
Phyllophaga saylori Sanderson, 1965
Phyllophaga schaefferi Saylor, 1937
Phyllophaga schenklingi Moser, 1918
Phyllophaga scitula (Horn, 1887)
Phyllophaga scoparia (LeConte, 1856)
Phyllophaga scuticeps (Bates, 1888)
Phyllophaga segregans (Bates, 1888)
Phyllophaga senex (Horn, 1878)
Phyllophaga sequoiana Saylor, 1936
Phyllophaga setifera (Burmeister, 1855)
Phyllophaga sinaloana Saylor, 1935
Phyllophaga sinuaticeps (Moser, 1921)
Phyllophaga skelleyi Woodruff and Beck, 1989
Phyllophaga snowi Saylor, 1940
Phyllophaga sociata (Horn, 1878)
Phyllophaga sodalis Reinhard, 1940
Phyllophaga sonora Saylor, 1939
Phyllophaga soror Davis, 1920
Phyllophaga spreta (Horn, 1887)
Phyllophaga squamipilosa Saylor, 1937
Phyllophaga stohleri Saylor, 1938
Phyllophaga submucida (LeConte, 1856)
Phyllophaga subpruinosa (Casey, 1884)
Phyllophaga subtonsa (LeConte, 1856)
Phyllophaga sylvatica Sanderson, 1942
Phyllophaga tarsalis (Schaeffer, 1908)
Phyllophaga taxodii Langston, 1924
Phyllophaga tecta Cartwright, 1944
Phyllophaga temaxa Saylor, 1940
Phyllophaga temora Saylor, 1943
Phyllophaga texensis Saylor, 1940
Phyllophaga timida (Horn, 1878)
Phyllophaga torta (LeConte, 1856)
Phyllophaga trichodes (Bates, 1890)
Phyllophaga tristis (Fabricius, 1781)
Phyllophaga trochanter Saylor, 1940
Phyllophaga tusa (Horn, 1887)
Phyllophaga ulkei (Smith, 1889)
Phyllophaga uniformis (Blanchard, 1851)
Phyllophaga vehemens (Horn, 1887)
Phyllophaga venodiola Saylor, 1948
Phyllophaga verruciventris (Moser, 1918)
Phyllophaga vetula (Horn, 1887)
Phyllophaga vexata (Horn, 1885)
Phyllophaga vilifrons (LeConte, 1856)
Phyllophaga wickhami Saylor, 1940
Phyllophaga xanthe (Bates, 1888)
Phyllophaga xerophila Saylor, 1939
Phyllophaga yaqui Saylor, 1940
Phyllophaga yemasseei Cartwright, 1944
Phyllophaga youngi Cartwright, 1935
Phyllophaga zavalana Reinhard, 1946
Planolinellus vittatus (Say, 1825)
Planolinoides aenictus (Cooper and Gordon, 1987)
Planolinoides borealis (Gyllenhal, 1827)
Planolinoides duplex (LeConte, 1878)
Planolinoides neotomae (Fall, 1907)
Planolinoides pectoralis (LeConte, 1857)
Planolinus tenellus (Say, 1823)
Platytomus atlanticus (Cartwright, 1948)
Platytomus caelicollis (Cartwright, 1948)
Platytomus longulus (Cartwright, 1948)
Platytomus micros (Bates, 1887)
Platytomus notialis (Cartwright, 1948)
Platytomus parvulus (Chevrolat, 1864)
Platytomus tibialis (Fabricius, 1798)
Plectris aliena Chapin, 1934
Plectrodes pubescens Horn, 1867
Plesiosternus setosus Morón, 1983
Pleurophorus caesus (Panzer, 1796)
Pleurophorus micros (Bates, 1887)
Pleurophorus parvulus (Chevrolat, 1864)
Podolasia emarginata Howden, 1954
Podolasia ferruginea (LeConte, 1856)
Podolasia involucris Howden, 1998
Podolasia lavignei Howden, 1997
Podolasia longipennis Howden, 1997
Podolasia peninsularis Howden, 1954
Podolasia pilosa Howden, 1954
Podolasia rotundipenis Howden, 1997
Podolasia stillwellorum Howden, 1997
Podolasia varicolor Saylor, 1948
Podostena bottimeri (Howden, 1958)
Podostena litoralis Howden, 1997
Podostena rileyi Howden, 1997
Podostena sleeperi Howden, 1997
Polyphylla aeola La Rue, 1998
Polyphylla anteronivea Hardy, 1978
Polyphylla arguta Casey, 1914
Polyphylla avittata Hardy, 1978
Polyphylla barbata Cazier, 1938
Polyphylla brownae Young, 1986
Polyphylla cavifrons LeConte, 1854
Polyphylla comes Casey, 1914
Polyphylla crinita LeConte, 1856
Polyphylla decemlineata (Say, 1824)
Polyphylla devestiva Young, 1966
Polyphylla diffracta Casey, 1891
Polyphylla donaldsoni Skelley, 2005
Polyphylla erratica Hardy, 1978
Polyphylla gracilis Horn, 1881
Polyphylla hammondi LeConte, 1856
Polyphylla hirsuta Van Dyke, 1933
Polyphylla mescalerensis Young, 1988
Polyphylla modulata Casey, 1914
Polyphylla monahansensis Hardy, 1978
Polyphylla multimaculata Hardy, 1981
Polyphylla nigra Casey, 1914
Polyphylla nubila Van Dyke, 1947
Polyphylla occidentalis (Linnaeus, 1767)
Polyphylla petitii (Guérin-Méneville, 1844)
Polyphylla pottsorum Hardy, 1978
Polyphylla pubescens Cartwright, 1939
Polyphylla ratcliffei Young, 1986
Polyphylla rugosipennis Casey, 1914
Polyphylla sobrina Casey, 1914
Polyphylla squamiventris Cazier, 1939
Polyphylla stellata Young, 1986
Polyphylla variolosa (Hentz, 1830)
Polyphylla woodruffi Skelley, 2005
Popillia japonica Newman, 1841
Protaetia fusca (Herbst, 1790)
Protaetia pryeri (Janson, 1888)
Psammodius basalis (Mulsant and Rey, 1870)
Psammodius laevipennis Costa, 1844
Psammodius pierottii Pittino, 1979
Pseudagolius bicolor (Say, 1823)
Pseudagolius coloradensis (Horn, 1870)
Pseudagolius warneri Gordon and Skelley, 2007
Pseudataenius contortus Cartwright, 1974
Pseudataenius socialis (Horn, 1871)
Pseudocanthon perplexus (LeConte, 1847)
Pseudocotalpa andrewsi Hardy, 1971
Pseudocotalpa giulianii Hardy, 1974
Pseudocotalpa sonorica Hardy, 1974
Psilocnemis leucosticta Burmeister, 1842
Rhysothorax rufus (Fabricius, 1792)
Rhyssemus brownwoodi Gordon and Cartwright, 1980
Rhyssemus californicus Horn, 1871
Rhyssemus germanus (Linnaeus, 1767)
Rhyssemus neglectus Brown, 1929
Rhyssemus scaber Haldeman, 1848
Rhyssemus sonatus LeConte, 1881
Rugaphodius rugatus (Schmidt, 1907)
Rutela formosa Burmeister, 1844
Rutelisca durangoana Ohaus, 1905
Saprosites pygmaeus Harold, 1877
Scabrostomus baileyi (Skelley and Gordon, 2002)
Scabrostomus bakeri (Skelley and Gordon, 2002)
Scabrostomus dyspistus (Skelley and Woodruff, 1991)
Scabrostomus kerni Gordon and Skelley, 2007
Scabrostomus parapeculiosus Gordon and Skelley, 2007
Scabrostomus peculiosus (Schmidt, 1922)
Scabrostomus sepultus (Cartwright, 1944)
Scabrostomus siccatus Gordon and Skelley, 2007
Scabrostomus skillmani (Warner and Skelley, 2006)
Scabrostomus tanytarsus (Skelley and Woodruff, 1991)
Schaefferellus arizonensis (Schaeffer, 1907)
Serica abdita Dawson, 1921
Serica acicula Dawson, 1932
Serica acontia Dawson, 1933
Serica adversa Dawson, 1967
Serica aemula Dawson, 1947
Serica alabama Dawson, 1967
Serica alleni Saylor, 1939
Serica alternata LeConte, 1856
Serica ammomenisco Hardy, 1987
Serica anthracina LeConte, 1856
Serica apatela Dawson, 1922
Serica arkansana Dawson, 1947
Serica aspera Dawson, 1922
Serica atracapilla (Kirby, 1837)
Serica atratula LeConte, 1856
Serica aviceps Dawson, 1967
Serica barri Dawson, 1967
Serica blatchleyi Dawson, 1932
Serica bruneri Dawson, 1967
Serica caliginosa Dawson, 1932
Serica campestris Dawson, 1919
Serica carolina Dawson, 1920
Serica catalina Dawson, 1947
Serica chaetosoma Dawson, 1932
Serica chicoensis Saylor, 1939
Serica coalinga Dawson, 1952
Serica concinna Dawson, 1947
Serica contorta Dawson, 1947
Serica craighead Saylor, 1939
Serica cruzi Saylor, 1939
Serica curvata LeConte, 1856
Serica cuyamaca Saylor, 1939
Serica delicata Dawson, 1922
Serica deserticola Dawson, 1952
Serica diablo Dawson, 1967
Serica egregia Dawson, 1921
Serica elmontea Saylor, 1939
Serica elongatula Horn, 1870
Serica elusa Dawson, 1919
Serica ensenada Saylor, 1948
Serica evidens Blatchley, 1919
Serica falcata Dawson, 1933
Serica falli Dawson, 1932
Serica fimbriata LeConte, 1856
Serica floridana Dawson, 1967
Serica frosti Dawson, 1967
Serica georgiana Leng, 1911
Serica heteracantha Dawson, 1967
Serica howdeni Dawson, 1967
Serica humboldti Gordon, 1975
Serica imitans Chapin, 1931
Serica intermixta Blatchley, 1910
Serica iricolor (Say, 1824)
Serica laguna Saylor, 1935
Serica ligulata Dawson, 1932
Serica lodingi Dawson, 1952
Serica loxia Dawson, 1920
Serica mckenziei Saylor, 1935
Serica mixta LeConte, 1856
Serica mystaca Dawson, 1922
Serica ochrosoma Dawson, 1919
Serica oliver Saylor, 1939
Serica opposita Dawson, 1921
Serica panda Dawson, 1952
Serica parallela Casey, 1884
Serica parvula (Blanchard, 1850)
Serica pavonia Dawson, 1932
Serica peleca Dawson, 1952
Serica perigonia Dawson, 1920
Serica pilifera Horn, 1894
Serica porcula Casey, 1884
Serica prava Dawson, 1933
Serica prunipennis Saylor, 1936
Serica psammobuna Hardy, 1987
Serica pullata Dawson, 1967
Serica pusilla Dawson, 1922
Serica repanda Dawson, 1933
Serica rhypha Dawson, 1952
Serica rossi Saylor, 1948
Serica sandiegensis Saylor, 1939
Serica satrapa Dawson, 1947
Serica scaphia Dawson, 1952
Serica sculptilis Dawson, 1922
Serica senta Dawson, 1933
Serica serensia Saylor, 1948
Serica sericea (Illiger, 1802)
Serica sericeoides Dawson, 1967
Serica serotina LeConte, 1856
Serica solita Dawson, 1922
Serica spicula Dawson, 1921
Serica sponsa Dawson, 1919
Serica stygia Dawson, 1933
Serica subnisa Dawson, 1947
Serica tantula Dawson, 1922
Serica texana LeConte, 1856
Serica tristis LeConte, 1850
Serica trociformis Burmeister, 1844
Serica ventura Dawson, 1932
Serica vespertina (Gyllenhal, 1817)
Serica watson Saylor, 1939
Setodius bryanti (Brown, 1929)
Setodius diogenes Gordon and Skelley, 2007
Setodius edmontonus (Brown, 1929)
Sisyphus spinipes Thunberg, 1818
Sisyphus submonticolus Howden, 1965
Stenotothorax badipes (Melsheimer, 1845)
Stenotothorax cadaverinus (Mannerheim, 1843)
Stenotothorax comosus Gordon and Skelley, 2007
Stenotothorax cribratulus (Schmidt, 1922)
Stenotothorax dilaticollis (Saylor, 1935)
Stenotothorax gardneri (Gordon, 2006)
Stenotothorax gentilis (Horn, 1887)
Stenotothorax gravis (Fall, 1927)
Stenotothorax lanei (Saylor, 1940)
Stenotothorax linsleyi (Van Dyke, 1933)
Stenotothorax martini (Van Dyke, 1928)
Stenotothorax mcpeaki Gordon and Skelley, 2007
Stenotothorax micellus Gordon and Skelley, 2007
Stenotothorax nevadensis (Horn, 1870)
Stenotothorax oriens Gordon and Skelley, 2007
Stenotothorax oviformis (Robinson, 1946)
Stenotothorax ovipennis (Horn, 1870)
Stenotothorax parapyriformis Gordon and Skelley, 2007
Stenotothorax pyriformis (Brown, 1929)
Stenotothorax rugoclypeus (Hinton, 1934)
Stenotothorax sparsus (LeConte, 1878)
Stenotothorax washtucna (Robinson, 1938)
Stenotothorax woodleyi Gordon, 2006
Strategus aloeus (Linnaeus, 1758)
Strategus antaeus (Drury, 1773)
Strategus cessus LeConte, 1866
Strategus craigi Ratcliffe, 1976
Strategus howdeni Ratcliffe, 1976
Strategus longichomperus Ratcliffe, 1976
Strategus mormon Burmeister, 1847
Strategus splendens (Palisot de Beauvois, 1809)
Strategus temoltzin Morón and Nogueira, 2008
Strigoderma arbicola (Fabricius, 1793)
Strigoderma costulipennis Bates, 1888
Strigoderma knausi (Brown, 1925)
Strigoderma pimalis Casey, 1884
Strigoderma protea Burmeister, 1844
Strigoderma pygmaea (Fabricius, 1798)
Strigoderma sulcipennis Burmeister, 1844
Strigoderma teapensis Bates, 1888
Strigoderma vestita Burmeister, 1844
Strigodius robinsoni (Cartwright, 1939)
Tesarius caelatus (LeConte, 1857)
Tesarius doyeni (Cartwright, 1977)
Tesarius mcclayi (Cartwright, 1955)
Tesarius oregonensis (Cartwright, 1955)
Tetraclipeoides acutissimus (Gordon, 1976)
Tetraclipeoides aemulus (Horn, 1887)
Tetraclipeoides anomaliceps (Brown, 1929)
Tetraclipeoides denticulatus (Haldeman, 1848)
Tetraclipeoides dentiger (LeConte, 1858)
Tetraclipeoides dentigerulus (Brown, 1929)
Tetraclipeoides fimbripes (Brown, 1928)
Tetraclipeoides formidatus (Gordon, 1976)
Tetraclipeoides giulianii (Gordon, 1977)
Tetraclipeoides henryi (Gordon, 1976)
Tetraclipeoides moquinus (Fall, 1907)
Tetraclipeoides testaceiventris (Fall, 1932)
Tetraclipeoides texanus Gordon and Skelley, 2007
Teuchestes fossor (Linnaeus, 1758)
Thyce deserta Hardy, 1974
Thyce squamicollis LeConte, 1856
Tomarus cuniculus (Fabricius, 1801)
Tomarus gibbosus (De Geer, 1774)
Tomarus laevicollis (Bates, 1888)
Tomarus nasutus (Burmeister, 1847)
Tomarus neglectus (LeConte, 1847)
Tomarus relictus (Say, 1825)
Tomarus sallaei (Bates, 1888)
Tomarus selanderi (Cartwright, 1959)
Tomarus subtropicus (Blatchley, 1922)
Trichiorhyssemus alternatus Hinton, 1938
Trichiorhyssemus cristatellus (Bates, 1887)
Trichiorhyssemus riparius (Horn, 1871)
Trichiotinus affinis (Gory and Percheron, 1833)
Trichiotinus assimilis (Kirby, 1837)
Trichiotinus bibens (Fabricius, 1775)
Trichiotinus lunulatus (Fabricius, 1775)
Trichiotinus piger (Fabricius, 1775)
Trichiotinus rufobrunneus (Casey, 1914)
Trichiotinus texanus (Horn, 1876)
Trichiotinus viridans (Kirby, 1837)
Trichonotulus scrofa (Fabricius, 1787)
Trigonopeltastes archimedes Schaum, 1841
Trigonopeltastes delta (Forster, 1771)
Trigonopeltastes discrepans Howden, 1968
Trigonopeltastes floridanus (Casey, 1909)
Trigonopeltastes frontalis Bates, 1889
Trigonopeltastes sallaei Bates, 1889
Trigonopeltastes truncatus Howden, 1968
Uroxys deavilai Delgado and Kohlmann, 2007
Valgus californicus Horn, 1870
Valgus canaliculatus (Olivier, 1789)
Valgus hemipterus (Linnaeus, 1758)
Valgus seticollis (Palisot de Beauvois, 1805)
Warwickia pilosa (Sanderson, 1939)
Xeropsamobeus acerbus (Horn, 1887)
Xeropsamobeus ambiguus (Fall, 1907)
Xeropsamobeus arenicolus Gordon and Skelley, 2007
Xeropsamobeus asellus (Schmidt, 1907)
Xeropsamobeus brighti Gordon and Skelley, 2007
Xeropsamobeus desertus (Van Dyke, 1918)
Xeropsamobeus doyeni Gordon and Skelley, 2007
Xeropsamobeus mohavei Gordon and Skelley, 2007
Xeropsamobeus padrei Gordon and Skelley, 2007
Xeropsamobeus scabriceps (LeConte, 1878)
Xyloryctes ensifer Bates, 1888
Xyloryctes jamaicensis (Drury, 1773)
Xyloryctes telephus Burmeister, 1847
Xyloryctes thestalus Bates, 1888

Подврсте

Anomala cincta cincta Say, 1835
Anomala flavilla coachellae Potts, 1977
Anomala flavilla flavilla Bates, 1888
Anomala flavipennis aransas Potts, 1977
Anomala flavipennis flavipennis Burmeister, 1844
Anomala flavipennis okaloosensis Potts, 1977
Anomala flavipennis subquadrata Casey, 1915
Canthon humectus alvarengai Halffter, 1961
Canthon humectus blumei Halffter and Halffter, 2003
Canthon humectus hidalgoensis Bates, 1887
Canthon humectus humectus (Say, 1832)
Copris klugi sierrensis Matthews, 1962
Copris lecontei lecontei Matthews, 1962
Copris remotus dicyrtus Matthews and Halffter, 1959
Copris remotus remotus LeConte, 1866
Cremastocheilus armatus armatus Walker, 1866
Cremastocheilus armatus cribripennis Casey, 1915
Cremastocheilus armatus maritimus Casey, 1915
Cremastocheilus armatus montanus Casey, 1915
Cremastocheilus castaneae brevisetosus Casey, 1915
Cremastocheilus castaneae castaneae Knoch, 1801
Cremastocheilus castaneae lecontei Westwood, 1874
Cremastocheilus castaneae pocularis Casey, 1915
Cremastocheilus crinitus bifoveatus Van Dyke, 1918
Cremastocheilus crinitus crinitus LeConte, 1874
Cremastocheilus crinitus pugetanus Casey, 1915
Cremastocheilus retractus incisus Casey, 1915
Cremastocheilus retractus retractus LeConte, 1874
Cremastocheilus schaumii schaumii LeConte, 1853
Cremastocheilus schaumii tibialis Casey, 1915
Cremastocheilus westwoodi tridens Casey, 1915
Cremastocheilus westwoodi westwoodi Horn, 1880
Cyclocephala lurida coahuilae Bates, 1888
Cyclocephala lurida lurida Bland, 1863
Deltochilum gibbosum gibbosum (Fabricius, 1775)
Deltochilum scabriusculum montanum Howden, 1966
Deltochilum scabriusculum scabriusculum Bates, 1887
Diplotaxis cribulosa cribulosa LeConte, 1856
Diplotaxis cribulosa sinaloa Vaurie, 1958
Diplotaxis moerens moerens LeConte, 1856
Diplotaxis moerens peninsularis Fall, 1909
Euphoria fascifera fascifera (LeConte, 1861)
Euphoria fascifera trapezium Casey, 1915
Euphoria fulgida fulgida (Fabricius, 1775)
Euphoria fulgida fuscocyanea Casey, 1915
Euphoria fulgida holochloris Fall, 1905
Euphoria fulgida limbalis Fall, 1905
Euphoria sepulchralis leucographa (Gory and Percheron, 1833)
Euphoria sepulchralis nitens Casey, 1915
Euphoria sepulchralis rufina (Gory and Percheron, 1833)
Euphoria sepulchralis sepulchralis (Fabricius, 1801)
Inca clathrata sommeri Westwood, 1845
Maladera castanea castanea (Arrow, 1913)
Malagoniella astyanax yucateca (Harold, 1863)
Onthophagus chevrolati chevrolati Harold, 1869
Onthophagus chevrolati longecarinatus Zunino and Halffter, 1988
Onthophagus fuscus fuscus Boucomont, 1932
Onthophagus hecate blatchleyi Brown, 1929
Onthophagus hecate hecate (Panzer, 1794)
Onthophagus landolti landolti Harold, 1880
Onthophagus landolti texanus Schaeffer, 1914
Onthophagus orpheus canadensis (Fabricius, 1801)
Onthophagus orpheus orpheus (Panzer, 1794)
Onthophagus orpheus pseudorpheus Howden and Cartwright, 1963
Onthophagus polyphemi polyphemi Hubbard, 1894
Onthophagus polyphemi sparsisetosus Howden and Cartwright, 1963
Onthophagus striatulus floridanus Blatchley, 1928
Onthophagus striatulus striatulus (Palisot de Beauvois, 1809)
Phanaeus amethystinus amethystinus Harold, 1863
Phanaeus triangularis texensis Edmonds, 1994
Phanaeus triangularis triangularis (Say, 1823)
Phyllophaga ephilida ephilida (Say, 1825)
Phyllophaga ephilida virilis Reinhard, 1939
Phyllophaga fraterna fraterna Harris, 1842
Phyllophaga fraterna mississippiensis Davis, 1920
Phyllophaga hirticula comosa Davis, 1920
Phyllophaga hirticula hirticula (Knoch, 1801)
Phyllophaga marginalis insolita Cartwright, 1944
Phyllophaga marginalis marginalis (LeConte, 1856)
Phyllophaga parvidens hysteropyga Davis, 1920
Phyllophaga parvidens parvidens (LeConte, 1856)
Phyllophaga tristis amplicornis Reinhard, 1939
Phyllophaga tristis suttonana Reinhard, 1939
Phyllophaga tristis tristis (Fabricius, 1781)
Phyllophaga vexata vexata (Horn, 1885)
Serica alternata alternata LeConte, 1856
Serica alternata exolita Dawson, 1933
Serica alternata patruela Dawson, 1933
Serica atratula atratula LeConte, 1856
Serica atratula monita Dawson, 1947
Serica georgiana georgiana Leng, 1911
Serica georgiana lecontei Dawson, 1921
Serica ligulata ligulata Dawson, 1932
Serica ligulata praetermissa Dawson, 1932
Serica perigonia eremicola Dawson, 1967
Serica perigonia perigonia Dawson, 1920
Serica ventura dorsalis Dawson, 1952
Serica ventura personata Dawson, 1932
Serica ventura ventura Dawson, 1932
Serica vespertina accola Dawson, 1921
Serica vespertina vespertina (Gyllenhal, 1817)
Trigonopeltastes sallaei sinaloensis Howden, 1968
Xyloryctes thestalus borealis Endrödi, 1975
Xyloryctes thestalus thestalus Bates, 1888

Референце

^ ^а ^б ^в ^г „Scarabaeidae”. BioRaS. Архивирано из оригинала 06. 03. 2018. г. Приступљено 05. 03. 2018.
^ Frolov, A.V. "Subfamily Scarabaeinae: atlas of representatives of the tribes (Scarabaeidae)". Retrieved on 2. 8. 2007
^ ^а ^б bugguide.net Family Scarabaeidae - Scarab Beetles

Литература

RU Ehlers. Current and Future Use of Nematodes in Biocontrol: Practice and Commercial Aspects with Regard to Regulatory Policy Issues. Biocontrol Science and Technology Volume 6, Issue 3, 1996.

Спољашње везе

Подаци везани за чланак Scarabaeidae на Викиврстама
Фотографије разних врста из породице (Scarabaeidae) на flickr.com
Scarabaeidae breeding site Photos of various Cetonidae, Dynastidae, Euchiridae, Lucanidae and Trichinae]
Family SCARABAEIDAE
June Beetles, Family: Scarabaeidae - Diagnostic photographs
Scarab Beetle Research, Databases, and Links from Scarab Central at University of Nebraska State Museum
Bibliography of literature published on scarab beetles since 1 January 2001 (worldwide coverage; through 2005)
UNL Архивирано на сајту Wayback Machine (26. јун 2012) Generic Guide to New World Scarabaeidae

[BioRaS-1] а ^б ^в ^г „Scarabaeidae”. BioRaS. Архивирано из оригинала 06. 03. 2018. г. Приступљено 05. 03. 2018.

[2] Frolov, A.V. "Subfamily Scarabaeinae: atlas of representatives of the tribes (Scarabaeidae)". Retrieved on 2. 8. 2007

[bugguide-3] а ^б bugguide.net Family Scarabaeidae - Scarab Beetles

[1]

[2]

[3]